Refereed Publications

Pryer, K.M. and D.J. Hearn. . Evolution of leaf form in marsileaceous ferns: evidence for heterochrony. Evolution (in press).

Schneider, H., A.R. Smith, and K.M. Pryer. . Is morphology really at odds with molecules in estimating fern phylogeny? Systematic Botany (in press).

Schuettpelz, E., A.L. Grusz, M.D. Windham, and K.M. Pryer. 2008. The utility of nuclear gapCp in resolving polyploid fern origins. Systematic Botany (in press).

Hennequin, S., E. Schuettpelz, K.M. Pryer, A. Ebihara, and J.-Y. Dubuisson. 2008. Divergence times and the evolution of epiphytism in filmy ferns (Hymenophyllaceae) revisited. International Journal of Plant Sciences (in press).

Pryer, K.M. and E. Schuettpelz. 2008. Ferns. In S.B. Hedges and S. Kumar (eds.) Timetree of Life. Oxford University Press, New York. (in press).

Nagalingum, N.S., M.D. Nowak, and K.M. Pryer. 2008. Assessing phylogenetic relationships in extant heterosporous ferns (Salviniales), with a focus on Pilularia and Salvinia. Botanical Journal of the Linnaean Society (in press).

Rothfels, C.J., M.D. Windham, A.L. Grusz, G.J. Gastony, and K.M. Pryer. 2008. Toward a monophyletic Notholaena (Pteridaceae): Resolving patterns of evolutionary convergence in xeric-adapted ferns. Taxon 57: 1-13.

Schuettpelz, E. and K.M. Pryer. 2008. Fern Phylogeny. Chapter 15 in Ranker, T.A. and C.H. Haufler (eds.). The Biology and Evolution of Ferns and Lycophytes. Cambridge Univ. Press., pp. 397-418.

Smith A.R., K.M. Pryer, E. Schuettpelz, P. Korall, H. Schneider, and P.G. Wolf. 2008. Fern Classification. Chapter 16 in Ranker, T.A. and C.H. Haufler (eds.). The Biology and Evolution of Ferns and Lycophytes. Cambridge Univ. Press., pp. 419-469.

Metzgar, J.S., J.E. Skog, E.A. Zimmer, and K.M. Pryer. 2008. The paraphyly of Osmunda is confirmed by phylogenetic analyses of seven plastid loci. Systematic Botany 33: 31-36.
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Christenhusz, M.J.M., H. Tuomisto, J.S. Metzgar, and K.M. Pryer. 2008. Evolutionary relationships within the Neotropical, eusporangiate fern genus Danaea (Marattiaceae). Molecular Phylogenetics and Evolution 46: 34-48.
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Schuettpelz E. and K.M. Pryer. 2007. Fern phylogeny inferred from 400 leptosporangiate species and three plastid genes. Taxon 56: 1037-1050.
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Haugen, P., D. Bhattacharya, J.D. Palmer, S. Turner, L.A. Lewis, and K.M. Pryer. 2007. Cyanobacterial ribosomal RNA genes with multiple, endonuclease-encoding group I introns. BMC Evolutionary Biology 7: 159.
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Metzgar, J.S., H. Schneider, and K.M. Pryer. 2007. Phylogeny and divergence time estimates for the fern genus Azolla (Salviniaceae). International Journal of Plant Sciences 168: 1045-1053.
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Schuettpelz E., H. Schneider, L. Huiet, M.D. Windham, and K.M. Pryer. 2007. A molecular phylogeny of the fern family Pteridaceae: assessing overall relationships and the affinities of previously unsampled genera. Molecular Phylogenetics and Evolution 44: 1172-1185.
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Korall, P., D.S. Conant, J.S. Metzgar, H. Schneider, K.M. Pryer. 2007. A molecular phylogeny of scaly tree ferns (Cyatheaceae). American Journal of Botany 94: 873-886 (cover).
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Nagalingum, N.S., H. Schneider, and K.M. Pryer. 2007. Molecular phylogenetic relationships and morphological evolution in the heterosporous fern genus Marsilea. Systematic Botany 32: 16-25.
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Schuettpelz, E., P. Korall, and K.M. Pryer. 2006. Plastid atpA data provide improved support for deep relationships among ferns. Taxon 55: 897-906.
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Schuettpelz, E. and K.M. Pryer. 2006. Reconciling extreme branch length differences: decoupling time and rate through the evolutionary history of filmy ferns. Systematic Biology 55: 485-502. [2006 Publisher's Award for Excellence in Systematic Research].
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Korall, P., K.M. Pryer, J. Metzgar, H. Schneider, D.S. Conant. 2006. Tree ferns: monophyletic groups and their relationships as revealed by four protein-coding plastid loci. Molecular Phylogenetics and Evolution 39: 830-845.
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Nagalingum, N.S., H. Schneider, and K.M. Pryer. 2006. Comparative morphology of reproductive structures in heterosporous water ferns and a reevaluation of the sporocarp. International Journal of Plant Sciences 167: 805-815.
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Korall, P., D.S. Conant, H. Schneider, K. Ueda, H. Nishida, K.M. Pryer. 2006. On the phylogenetic position of Cystodium: it’s not a tree fern – it's a polypod! American Fern Journal 96: 45-53.
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Smith, A.R., K.M. Pryer, E. Schuettpelz, P. Korall, H. Schneider, and P.G. Wolf. 2006. A classification for extant ferns. Taxon 55: 705-731.
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Wikström, N. and K.M. Pryer. 2005. Incongruence between primary sequence data and the distribution of a mitochondrial atp1 group II intron among ferns and horsetails. Molecular Phylogenetics and Evolution 36: 484-493.
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Schneider, H., E. Schuettpelz, K.M. Pryer, R. Cranfill, S. Magallón, R. Lupia. 2004. Ferns diversified in the shadow of angiosperms. Nature 428: 553-557.
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Pryer, K.M., H. Schneider and S. Magallón. 2004. The radiation of vascular plants. In J. Cracraft and M. J. Donoghue, eds. Assembling the Tree of Life, 138-153. Oxford University Press, London

Pryer, K.M., E. Schuettpelz, P.G. Wolf, H. Schneider, A.R. Smith, R. Cranfill. 2004. Phylogeny and evolution of ferns (monilophytes) with a focus on the early leptosporangiate divergences. American Journal of Botany 91: 1582-1598. [Special invited paper]
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Des Marais, D.L., A.R. Smith, D.M. Britton, and K.M. Pryer. 2003. Phylogenetic relationships and evolution of extant horsetails, Equisetum, based on chloroplast DNA sequence data (rbcL and trnL-F). International Journal of Plant Sciences 164 (5): 737-751.
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Dubuisson, J.-Y., S. Hennequin, E.J.P. Douzery, R. Cranfill, A.R. Smith and K.M. Pryer. 2003. rbcL phylogeny of the fern genus Trichomanes (Hymenophyllaceae), with special reference to neotropical taxa. International Journal of Plant Sciences 164 (5): 753-761.
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Pryer, K.M., H. Schneider, E.A. Zimmer and J.A. Banks. 2002. Deciding among green plants for whole genome studies. Trends in Plant Science 7: 550-554.
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Schneider, H. and K.M. Pryer. 2002. Structure and function of spores in the aquatic heterosporous fern family Marsileaceae. International Journal of Plant Sciences 163: 485-505 (cover).
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Schneider, H., K.M. Pryer, R. Cranfill, A.R. Smith and P.G. Wolf. 2002. Evolution of vascular plant body plans: a phylogenetic perspective. Pp. 330-364 in Developmental Genetics and Plant Evolution, Q.C.B. Cronk, R.M. Bateman, and J.A. Harris, eds. Taylor and Francis, Philadelphia.

Smith, A.R., H. Tuomisto, K.M. Pryer, J.S. Hunt and P.G. Wolf. 2001. Metaxya lanosa, a second species in the genus and fern family Metaxyaceae. Systematic Botany 26: 480-486.
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Pryer, K.M., A.R. Smith, J.S. Hunt, and J.-Y.Dubuisson. 2001. rbcL data reveal two monophyletic groups of filmy ferns (Filicopsida:Hymenophyllaceae). American Journal of Botany 88: 1118-1130 (cover).
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Pryer, K.M., H. Schneider, A.R. Smith, R. Cranfill, P.G. Wolf, J.S. Hunt and S.D. Sipes. 2001. Horsetails and ferns are a monophyletic group and the closest living relatives to seed plants. Nature 409: 618-622 (cover).
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Lupia, R., H. Schneider, G.M. Moeser, K.M. Pryer and P.R. Crane. 2000. Marsileaceae sporocarps and spores from the Late Cretaceous of Georgia, U.S.A. International Journal of Plant Sciences 161: 975-988.
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Wolf, P.G., S.D. Sipes, M.R. White, M.L. Martines, K.M. Pryer, A.R. Smith and K. Ueda. 1999. Phylogenetic relationships of the enigmatic fern families Hymenophyllopsidaceae and Lophosoriaceae: evidence from rbcL nucleotide sequences. Plant Systematics and Evolution 219: 263-270.
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Turner, S., K.M. Pryer, V.P.W. Miao and J.D. Palmer. 1999. Investigating deep phylogenetic relationships among cyanobacteria and plastids by small subunit rRNA sequence analysis. Journal of Eukaryotic Microbiology 46: 327-338.
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Pryer, K.M. 1999. Phylogeny of marsileaceous ferns and relationships of the fossil Hydropteris pinnata reconsidered. International Journal of Plant Sciences 160:931-954 (cover).
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Wolf, P.G., K.M. Pryer, A.R. Smith and M. Hasebe. 1998. Phylogenetic studies of extant pteridophytes. Pp. 541-556 in D.E. Soltis, P.S. Soltis and J.J. Doyle, eds. Molecular systematics of plants II: DNA sequencing. Kluwer Academic Publishers, Boston.

Pryer, K.M., A.R. Smith and J.E. Skog. 1995. Phylogenetic relationships of extant ferns based on evidence from morphology and rbcL sequences. American Fern Journal 85 (Special Issue): 205-282.
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Hasebe, M., P.G. Wolf, K.M. Pryer, K. Ueda, M. Ito, R. Sano, G.J. Gastony, J. Yokoyama, J.R. Manhart, N. Murakami, E.H. Crane, C.H. Haufler and W.D. Hauk. 1995. Fern phylogeny based on rbcL nucleotide sequences. American Fern Journal 85 (Special Issue): 134-181.
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Pryer, K.M. and C.H. Haufler. 1993. Isozymic and chromosomal evidence for the allotetraploid origin of Gymnocarpium dryopteris (Dryopteridaceae). Systematic Botany 18: 150-172.
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Pryer, K.M. 1993. Gymnocarpium. Pp. 258-262 in Flora of North America North of Mexico, Vol.2. Flora of North America Editorial Committee (ed.). Oxford University Press, New York.
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Pryer, K.M. 1992. The status of Gymnocarpium heterosporum and G. robertianum in Pennsylvania. American Fern Journal 81: 34-39.

Pryer, K.M. 1990. The limestone oak fern:new to Manitoba. The Blue Jay 48 (4):192-195.
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Pryer, K.M. and L.R. Phillippe. 1989. A synopsis of the genus Sanicula in eastern Canada. Canadian Journal of Botany 67: 694-707.
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McNeill, J. and K.M. Pryer. 1985. The status and typification of Phegopteris and Gymnocarpium. Taxon 34: 136-156.

Pryer, K.M., D.M. Britton and J. McNeill. 1983. A numerical analysis of chromatographic profiles in North American taxa of the fern genus Gymnocarpium. Canadian Journal of Botany 61: 2592-2602.

Pryer, K.M. and D.M. Britton. 1983. Spore studies in the genus Gymnocarpium. Canadian Journal of Botany 61: 377-388.

Sarvela, J., D.M. Britton and K.M. Pryer. 1981. Studies on the Gymnocarpium robertianum complex in North America. Rhodora 83: 421-431.

Chase, R., K.M. Pryer, R. Baker and D.M. Madison. 1978. Responses to conspecific chemical stimuli in the terrestrial snail Achatina fulica (Pulmonata: Sigmuretha). Behavioural Biology 22: 302-315.

Books

Argus, G.W. & K.M. Pryer. 1990. Rare Vascular Plants in Canada - Our Natural Heritage. Canadian Museum of Nature, Ottawa. 277 pp.

Pryer, K.M. & G.W. Argus (eds.). 1987. Atlas of the Rare Vascular Plants of Ontario. Part 4. National Museum of Natural Sciences, Ottawa. 472 pp.

Electronic Publications

Pryer, K.M. and S.J.Hackett. 1999. Field's Museum's Pritzker Laboratory of Molecular Systematics and Evolution.
Web site

Pryer, K.M. 1999. Phylogeny,character evolution,and diversification of extant ferns.In addition to being a website created for NSF grant DEB-9615533, this site permits access to an online Fern DNA Database.
Web site
Selection for the Scout Report for Science & Engineering, November 24, 1999
HMS Beagle's "Web Pick of the Day", December 17, 1999

Pryer, K.M. and A.R. Smith. 1997. Filicopsida. In Maddison, D.R.and W.P.Maddison. The Tree of Life: A distributed Internet project containing information about phylogeny and biodiversity.
Web site

Pryer, K.M. and A.R. Smith. 1997. Leptosporangiate Ferns. In Maddison, D.R.and W.P. Maddison. The Tree of Life: A distributed Internet project containing information about phylogeny and biodiversity.
Web site